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Mostly because you don't want to have to go straight home from wherever you were using the gun, and then head back out to shop.
Had a roommate that routinely put a couple of guns in the trunk of his car during hunting season. He'd go out hunting in the morning before work, or stop off on the way home afterwards. One rifle and one handgun. Rifle for shooting the deer or elk, handgun for finishing off the animal. (If it was bird hunting season, he'd just have a shotgun. We both kept fishing tackle in our cars when going to work when fishing was legal, but lures and monofilament really aren't problems.)
I've been with groups that were out target shooting and on the way home they stopped for pizza or other activities. To drop off 10 people and their guns and then reassemble made little sense. Having 10 people carrying handguns and blackpowder rifles suddenly descend on a Dairy Queen or a pizza place would probably make the owners a bit nervous--then what do you do when you have to go pee, take them with you? I've known people who'd go out hunting and do the same: Into the trunk with them.
It's what my brother does on the way to and from the shooting range with his son. Trunks are better than extra trips or taking the guns in with you when buying ice and sodas.
Then there's the problematic hunting + camping trip. You go hunting for a weekend, and camp. When not in use, the safest place is the trunk of the car or locked in a box in the bed of a truck. That includes when you're shopping for food or grabbing a burger. The alternative--much safer, apparently--is leaving the guns "locked" in your tent. In all cases, serious hunters also had a handgun for finishing their prey.
Had another roommate who worked driving an armored car. Whenever there was a report of an armored car robbery, he'd go a bit paranoid. Granted, his employer also had no problem with his carrying guns--he'd bike to work in summer with one gun in a holster under his t-shirt and a couple more in a backpack, through a public park. After work he'd really have no choice but to bike straight home. But during the rest of the year he'd do errands after work, and keep them locked in the trunk of his car. Better than taking them into the grocery store, or the, er, novelty shops that he'd visit before heading to his girlfriend's for the night. Just imagine: "Is that a gun in your pocket, or do you like the magazine?" And he'd pull out his Glock. Now *that* would go over well. (At home he kept pretty much everything in a gun safe, although he said to be sure to make a lot of noise when coming home late at night--he assumed that a thief would be loud, but I wouldn't be, and he'd rather get woken up by a rude housemate than run the risk of having to find another housemate. He said he kept one next to his bed.)
I could probably go on, but you get the picture. Different sort of lifestyle, for some people, and for others, just part of work.
This doesn't include people that just want to feel safe and carry a gun in their car. As far as I know, I've never known anybody like that, but I guess they exist.
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